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This cheat sheet summarizes the major metabolic pathways that cells use to harvest, store, and redirect chemical energy. It connects carbohydrate, lipid, and amino acid metabolism so students can see how pathways feed into one another. College biochemistry requires tracking carbon flow, redox carriers, ATP yield, and regulation, so a compact pathway overview helps organize many details.

Use it to compare pathway locations, inputs, outputs, and control points.

Key Facts

  • Glycolysis converts one glucose to two pyruvate with net products 2ATP2\mathrm{ATP} and 2NADH2\mathrm{NADH}: glucose+2ADP+2Pi+2NAD+2pyruvate+2ATP+2NADH+2H++2H2O\mathrm{glucose} + 2\mathrm{ADP} + 2\mathrm{P_i} + 2\mathrm{NAD^+} \rightarrow 2\mathrm{pyruvate} + 2\mathrm{ATP} + 2\mathrm{NADH} + 2\mathrm{H^+} + 2\mathrm{H_2O}.
  • Pyruvate dehydrogenase links glycolysis to the TCA cycle by forming acetyl-CoA: pyruvate+CoA+NAD+acetyl-CoA+CO2+NADH+H+\mathrm{pyruvate} + \mathrm{CoA} + \mathrm{NAD^+} \rightarrow \mathrm{acetyl\text{-}CoA} + \mathrm{CO_2} + \mathrm{NADH} + \mathrm{H^+}.
  • Each acetyl-CoA oxidized in the TCA cycle yields 3NADH3\mathrm{NADH}, 1FADH21\mathrm{FADH_2}, 1GTP1\mathrm{GTP}, and 2CO22\mathrm{CO_2}.
  • Oxidative phosphorylation uses electrons from NADH\mathrm{NADH} and FADH2\mathrm{FADH_2} to drive proton pumping, and ATP synthase uses the gradient to make ATP from ADP+Pi\mathrm{ADP} + \mathrm{P_i}.
  • Approximate ATP yields are 2.5ATP2.5\mathrm{ATP} per NADH\mathrm{NADH} and 1.5ATP1.5\mathrm{ATP} per FADH2\mathrm{FADH_2} under typical mitochondrial conditions.
  • Beta-oxidation shortens a saturated fatty acyl-CoA by two carbons per cycle, producing 1FADH21\mathrm{FADH_2}, 1NADH1\mathrm{NADH}, and 1acetyl-CoA1\mathrm{acetyl\text{-}CoA} per cycle except for the final cleavage pattern.
  • Gluconeogenesis bypasses irreversible glycolysis steps using pyruvate carboxylase, PEP carboxykinase, fructose 1,61,6-bisphosphatase, and glucose 66-phosphatase.
  • Reciprocal regulation prevents futile cycling, so high ATP\mathrm{ATP} and citrate generally slow glycolysis while high AMP\mathrm{AMP} and fructose 2,62,6-bisphosphate promote glycolysis.

Vocabulary

Metabolic pathway
A linked series of enzyme-catalyzed reactions that transforms molecules through defined intermediates.
Catabolism
The set of pathways that break down molecules and often capture energy as ATP\mathrm{ATP}, NADH\mathrm{NADH}, or FADH2\mathrm{FADH_2}.
Anabolism
The set of pathways that build complex molecules and usually require energy input such as ATP\mathrm{ATP} or reducing power such as NADPH\mathrm{NADPH}.
Acetyl-CoA
A two-carbon activated carrier that delivers acetyl groups to the TCA cycle, lipid synthesis, ketone body formation, and other pathways.
Redox carrier
A molecule such as NAD+\mathrm{NAD^+}, NADP+\mathrm{NADP^+}, or FAD\mathrm{FAD} that transfers electrons during oxidation-reduction reactions.
Allosteric regulation
Control of enzyme activity when a molecule binds at a site other than the active site and changes enzyme shape or function.

Common Mistakes to Avoid

  • Counting gross ATP instead of net ATP in glycolysis is wrong because the pathway uses 2ATP2\mathrm{ATP} early and produces 4ATP4\mathrm{ATP} later, giving net 2ATP2\mathrm{ATP} per glucose.
  • Treating NADH\mathrm{NADH} and NADPH\mathrm{NADPH} as interchangeable is wrong because NADH\mathrm{NADH} mainly supports ATP production while NADPH\mathrm{NADPH} mainly supports biosynthesis and antioxidant defense.
  • Placing all metabolic pathways in the same compartment is wrong because glycolysis occurs in the cytosol, the TCA cycle occurs in the mitochondrial matrix, and electron transport occurs in the inner mitochondrial membrane.
  • Assuming gluconeogenesis is simply glycolysis in reverse is wrong because several glycolysis reactions are irreversible and require separate bypass enzymes.
  • Forgetting carbon loss as CO2\mathrm{CO_2} is wrong because carbon accounting is essential for understanding pyruvate oxidation, the TCA cycle, and amino acid entry points.

Practice Questions

  1. 1 One glucose molecule produces 2NADH2\mathrm{NADH} in glycolysis, 2NADH2\mathrm{NADH} from pyruvate dehydrogenase, and 6NADH6\mathrm{NADH} plus 2FADH22\mathrm{FADH_2} in the TCA cycle. Using 2.5ATP2.5\mathrm{ATP} per NADH\mathrm{NADH} and 1.5ATP1.5\mathrm{ATP} per FADH2\mathrm{FADH_2}, how many ATP equivalents come from these reduced carriers?
  2. 2 A saturated 1616-carbon fatty acyl-CoA undergoes beta-oxidation. How many acetyl-CoA molecules are produced, and how many beta-oxidation cycles occur?
  3. 3 If one turn of the TCA cycle produces 3NADH3\mathrm{NADH}, 1FADH21\mathrm{FADH_2}, and 1GTP1\mathrm{GTP}, estimate the ATP equivalent yield per acetyl-CoA using 2.5ATP2.5\mathrm{ATP} per NADH\mathrm{NADH}, 1.5ATP1.5\mathrm{ATP} per FADH2\mathrm{FADH_2}, and 1ATP1\mathrm{ATP} per GTP\mathrm{GTP}.
  4. 4 Explain why a cell should not run glycolysis and gluconeogenesis at high rates at the same time, and identify one regulatory molecule that helps prevent this.